| Student of Novosibirsk State University (Russia), Dep. of General Biology, Faculty of Natural Sciences | 1990 – 1995 |
| PhD student of Institute of Systematics and Ecology of Animals SB RAS (Laboratory of Insect Ecology) [Ph. D. thesis, 2002, theme: ‘The ethological and ecological aspects of ant-aphid interactions’], specialization: Entomology]. | 1998–2001 |
| Dissertation to obtain Doctor of science degree, specialization: Entomology. Theme: “Ecological and ethological aspects of interaction between ants, aphids and aphidophages at different levels of social organization.” Institute of Systematics and Ecology of Animals SB RAS, Novosibirsk. 306 p. | 2015 |
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